下列关于马进化过程的叙述中,符合达尔文学说的观点是()

A、广阔草原对始祖马进行了定向选择

B、始祖马的不定向变异,是因为环境的改变

C、马趾产生的变异,都能适应环境条件的改变

D、始祖马趾的变化,是因为生活环境由灌木丛到广阔的草原

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